Konrad Halupka [ha - woop -ca] konrad.halupka@uwr.edu.pl

assistant professor at the Department of Behavioural Ecology

konrad halupka

Research interests: behavioural ecology (animal communication, antipredator and foraging tactics), complex systems (spreading in networks, emergent behaviour), natural history and climate change, predicting the future

Potential areas for collaboration: statistical analysis (bootstrapping, resampling, meta-analyses), agent-based modelling, exciting field research

Citation report in Google Scholar


Halupka K. 1994. Incubation feeding in Meadow Pipit Anthus pratensis affects female time budget. Journal of Avian Biology 25: 251-253.

Halupka K. 1998. Nest predation in Meadow Pipits Anthus pratensis nesting in natural conditions. Ornis Fennica 75: 139-143.

Halupka K. 1998. Nest-site selection and nest predation in Meadow Pipits. Folia Zoologica 47: 29-37.

Halupka K. 1998. Partial nest predation in an altricial bird selects for the accelerated development of young. Journal of Avian Biology 29: 129-133.

Halupka K. 1998. Vocal begging by nestlings and vulnerability to nest predation in Meadow Pipits Anthus pratensis; to what extent do predation costs of begging exist? Ibis 140: 144- 149.

Halupka K. & Hałupka L. 1997. The influence of reproductive season stage on nest defence by meadow pipit (Anthus pratensis). Ethology Ecology & Evolution 9: 89-98.

Halupka K. & Hałupka L. 1998. Alarm calls and chick reaction: comments on Kleindorfer et al. (1996). Animal Behaviour 55: 502-503.

Halupka K., Borowiec M., Karczewska A., Kuńka A. & Pietrowiak J. 2002. Habitat requirements of Whitethroats Sylvia communis breeding in an alluvial plain. Bird Study 49: 297-299.

Hutchinson J.M.C., Halupka K. 2004. Mate choice when males are in patches: optimal strategies and good rules of thumb. Journal of Theoretical Biology 231: 129-151.

Wysocki D., Halupka K. 2004. The frequency and timing of courtship and copulation in blackbirds, Turdus merula, reflect sperm competition and sexual conflict. Behaviour 141: 501-512.

Greeney H.F., Dobbs R.C., Martin P. R., Halupka K., Gelis R. A. 2005. Nesting and foraging ecology of the rufous-crowned tody-tyrant (Poecilotricus ruficeps) in Eastern Ecuador. Ornitologia Neotropical 16: 427-432.

Halupka K., Borowiec M. 2006. Male whitethroats, Sylvia communis, advertise their future contribution to parental care. Behaviour 143: 1-14.

Dyrcz A., Halupka K. 2007. Why does the frequency of nest parasitism by the cuckoo differ considerably between two populations of warblers living in the same habitat? Ethology 113: 209-213.ethology

Borowiec M., Łosak K., Osiejuk T. S., Halupka K. 2007. Song repertoire is not affected by stress in an adult male songbird, the Whitethroat Sylvia communis. Behaviour 144: 879-887.ethology

Greeney H.F., Halupka K. 2008. Nesting biology of the Andean Solitaire (Myadestes ralloides) in Northeastern Ecuador. Ornitologia Neotropical 19: 213-219.

Halupka K., Greeney H.F. 2009. Breeding biology of Pale-eyed Thrushes (Turdus leucops) in the cloud forest of northeastern Ecuador. Ornitologia Neotropical 20: 381-389.

Halupka K., Greeney H.F. 2009. The influence of parental behaviour on vulnerability to nest predation in tropical thrushes of an Andean cloud forest. Journal of Avian Biology 40: 658-661.

Halupka K., Osińska-Dzienniak A. 2013. Group foraging by wintering rooks: combined effects of safety and energy requirements. Behaviour 150: 381-395.

Halupka K. 2014. Spreading information in a network of interacting neighbours. PLoS ONE 9: e102801.

Hałupka L., Halupka K., Klimczuk E., Sztwiertnia H. 2014. Coping with shifting nest predation refuges by European reed warblers Acrocephalus scirpaceus. PLoS ONE 9: e115456.

Halupka K. 2017. Information flow in a network of dispersed signalers-receivers. Physica A 486: 610-617.

Hałupka L., Halupka K. 2017. The effect of climate change on the duration of avian breeding seasons: a meta-analysis. Proceedings of the Royal Society B 284: 20171710.

Hałupka L., Sztwiertnia H., Marczuk M., Dziachan A., Kosmowska A., Klimczuk E., Halupka K. 2018. Ageing nestlings of the Reed Warbler Acrocephalus scirpaceus. Ringing & Migration 33: 1-9.

Halupka K., Błaszkowska J., Kołodziejczyk N., Kulik D., Nowak P., Przepiórka E., Siemianowska S. 2018. Error analysis in the estimation of coral reef fish numbers from visual surveys. Galaxea, Journal of Coral Reef Studies 20: 13-18.

Najbar A., Konowalik A., Halupka K., Najbar B., Ogielska M. 2020. Body size and life history traits of the fire salamander (Salamandra salamandra) from Poland. Amphibia-Reptalia 41: 63-74.

Halupka L., Borowiec M., Neubauer G., Halupka K. 2021. Fitness consequences of longer breeding seasons of a migratory passerine under changing climatic conditions. Journal of Animal Ecology; https://doi.org/10.1111/1365-2656.13481

For the general public

Halupka K. 1991 Niewierność małżeńska u ptaków. [Mixed reproductive strategy in birds] Przeglad Zoologiczny 35: 221-244.

Halupka K. 1992. Opieka rodzicielska u zwierząt. [Parental care in animals] Problemy 548: 27-33.

Halupka K. 2000. Drapieżnik doda ci skrzydeł. [A predator will add you wings] Wiedza i Zycie 785: 22-25.

Halupka K. 2004. Jak to jest z papugowaniem? [Do parrots parrot?] Woliera 15: 4-7.

Halupka K. 2007. Rospuda pod Newbury. Polityka 16: 113-115.


J_test - to compare two Mayfield's estimators of daily survival rates.

  • asks for number of nest-days (successful and with losses)

  • calculates Mayfield's statistics, their standard errors and 95% confidence intervals

  • compares two daily survival rates using the test developed by D.H. Johnson (1979. Estimating nest success: the Mayfield method and an alternative. Auk 96: 651 - 661)

  • displays the effect size (and its 95% c.l.), test statistics and P-value

download (windows)

source (FORTRAN 77)

If you need help...

Look up the original papers by Mayfield (1975) and Johnson (1979) available from the PRBO Conservation Science. Mayfield & Johnson combo is a reasonably good and robust method, escpecially when the sample size is small, but you have a "good" data (short intervals between nest checks).

Fortran routines for counting clusters and detecting spanning clusters in a regular, 8-neighbors network with boundaries

I work on signal transmission in a network of interacting neighbours, which pass signals between each other. I wrote Fortran programs which simulate and analyse regular networks, where each node has 8 nearest neighbors (or 5 or 3, when it is on the boundary), also known as the Moore neighborhood. Here are two subroutines: "klaster" counts number of clusters of active nodes and "spanning" (called from "klaster") detects if there is any top-down spanning (percolating) cluster in the network:

subroutines "klaster" and "spanning" (FORTRAN 77)

Three exemplary small lattices (5x5 nodes) are shown below (active nodes are red). The upper one has two clusters (with the biggest one including 7 nodes) and no spanning clusters. On the next graph there is only one and spanning cluster of size 13. The last one has two clusters and the biggest (and spanning) cluster has 11 nodes.

clusters in 8-neighbors networks

An exemplary program using "klaster" and "spanning" subs:


creates several networks with random pattern of active clusters, shows each network in the text mode and performs the analysis (here is an output from one run)

Update: Apr 2021

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